Cryptographic software needs a source of unpredictable data to work correctly. Many open source operating systems provide a "randomness device" that serves this purpose (usually named /dev/random). On other systems, applications have to seed the OpenSSL Pseudo Random Number Generator (PRNG) manually with appropriate data before generating keys or performing public key encryption.
As of version 0.9.5, the OpenSSL functions that need randomness report an error if the PRNG has not been seeded with at least 128 bits of randomness. So mod_ssl has to provide enough entropy to the PRNG to work correctly. For this one has to use the SSLRandomSeed directives.
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