Why is there still white space between these divs in all browsers except Chrome?

It's probably because your images are using the default display: inline property which applies line-height to the image, usually resulting in a bit of whitespace at the bottom of the image.

For some reason, the #btnWrapper div was forced to have a greater line-height than necessary in IE and FF. Adding a line-height of (at most) .7em proved to close up the white space (kind of funny because the actual buttons themselves (which are contained within the #btnWrapper div) are rendered with a line height of 1.3em, so this makes no real sense to me, but...).

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